Abstract
Pygmy grasshoppers (Tetrigidae) are a speciose group of complicated taxonomy, with many species requiring clarification on their species boundaries, and more still awaiting discovery. Two new species of Scelimena Serville, 1838 are described: (1) from S. discalis species group S. gombakensis sp. nov. and (2) from S. hexodon species group Scelimena marta sp. nov. Catalogue of Scelimenini genera (15) and species (100) is presented and taxonomy and biogeography of the tribe are discussed. New and resurrected combinations are: Falconius becvari (Buzzetti & Devriese, 2008) comb. nov. (of Gavialidium becvari), Gavialidium carli Hebard, 1930 comb. resurr. (of Bidentatettix carli), Indoscelimena india (Hancock, 1907) comb. nov. (of Scelimena india), Paragavialidium nodiferum (Walker, 1871) comb. nov. (of Platygavialidium nodiferum), Platygavialidium productum (Walker, 1871) comb. nov. (of Gavialidium productum), Scelimena hexodon (Haan, 1843) comb. resurr. (of Hexocera hexodon), Scelimena rosacea (Hancock, 1915) comb. resurr. (of Amphibotettix rosaceus), Tegotettix bufocrocodil (Storozhenko & Dawwrueng, 2015) comb. nov. (of Gavialidium bufocrocodil). New synonyms are: Gavialidium phangensum Mahmood, Idris & Salmah, 2007 syn. nov. (of Eufalconius pendleburyi), Gavialidium philippinum Bolívar, 1887 syn. nov. (of Platygavialidium productum comb. nov.), Hexocera Hancock, 1915 syn. nov. (of Scelimena), Paracriotettix Liang, 2002 syn. nov. (of Scelimena), Paracriotettix zhengi Liang, 2002 syn. nov. (of Scelimena melli), Scelimena mellioides Deng, 2016 syn. nov. (of Scelimena melli), Scelimena wuyishanensis Deng, 2016 syn. nov. (of Platygavialidium sinicum). The genus Scelimena is divided into six species groups. Eucriotettix neesoon Tan & Storozhenko nom. nov. is new name for homonym Eucriotettix guentheri Tan & Storozhenko, 2017. Finally, a tabular key to 15 Scelimenini genera, based on 16 morphological characters, is presented.
References
Bey-Bienko, G. & Mistshenko, L. (1951) Keys to the Fauna of the U.S.S.R. [1964 English translation No. 40]. In: Bey-Bienko, G. & Mistshenko, L. (Eds.) Locusts and Grasshoppers of the U.S.S.R. and Adjacent Countries. Vol. 2. Academy of Sciences, Moscow/Leningrad, pp. 385–667.
Bhalerao, A.M. & Paranjape, S.Y. (1986) Studies on the bioecology of a grouse locust Euscelimena harpago Serv. (Orthoptera: Tetrigidae). Gegbios, 13, 145–150.
Bhalerao, A.M., Naidu N.M. & Paranjape, S.Y. (1987) Some observations on the nutrition—reproduction correlation in grouse locusts (Orthoptera: Tetrigidae). Proceedings of the Indian Academy of Sciences (Animal Sciences), 96 (3), 323–327.
https://doi.org/10.1007/BF03180016Blackith, R.E. (1988) The Tetrigidae (Orthoptera) of Sri Lanka. Entomologica Scandinavica, 30 (Supplement), 93–109.
Blackith, R.E. (1992) The Tetrigidae (Insecta; Orthoptera) of South-East Asia; an Annotated Catalogue with Partial Translated Keys and Bibliography. Japaga Press, Dublin, LIV + 249 pp.
Bolívar, I. (1887) Essai sur les Acridiens de la tribu des Tettigidae. Annales de la Sociét Entomologique de Belgique, 31, 175–313, pls. 4–5.
Bolívar, I. (1898) Contributions l'étude des Acridiens espèces de la Faune indo et austro-malaisienne du Museo Civico di Storia Naturale di Genova. Annali del Museo Civico di Storia Naturale di Genova, 39, 66–101.
https://doi.org/10.5962/bhl.part.9541Bolívar, I. (1902) Ortópteros nuevos de España. Boletín de la Real Sociedad Española de Historia Natural, 2, 86–88.
Bolívar, I. (1906) Fasgonurídeos de la Guinea Española. Memorias de la Real Sociedad Española de Historia Natural, 1, 327–377.
Bolívar, I. (1909) Nouvelles espèces d'Acridiens du Musée de Genève. Boletín de la Real Sociedad Española de Historia Natural, 9, 393–403.
Bolívar, I. (1917[1918]) Contribución al conocimiento de la fauna índica. Orthoptera (Locustidae vel Acridiidae). Revista de la Real Academia de Ciencias Exactas, Físicas y Naturales, 16, 278–289, 374–412.
Bolívar, I. (1918) Estudios entomológicos. Tercera parte. Sección Oxyae. Trabajos del Museo de Ciencias Naturales (Serie Zoológica), 34, 1–43.
Bruner, L. (1915) Preliminary catalogue of the Orthopteroid insects of the Philippine Islands. University Studies, 15 (2), 195–281. [1–87]
Brunner von Wattenwyl, K. (1893) Révision du système des Orthoptères et déscription des espèces rapportées par M. Leonardo Fea de Birmanie. Annali del Museo Civico di Storia Naturale di Genova, 13 (33), 1–230.
https://doi.org/10.5962/bhl.title.5121Buzzetti, F.M. & Devriese, H. (2008) On some Oriental Orthoptera, mostly from Myanmar (Insecta: Orthoptera: Ensifera, Caelifera). Bollettino del Museo Civico Storia naturale di Verona, 32, 161–169.
Cigliano, M.M., Braun H., Eades D.C. & Otte D. (2018) Orthoptera Species File. Version 5.0/5.0. Available from: http://Orthoptera.SpeciesFile.org (accessed 14 August 2018)
Casto de Elera, R.P.F (1895) Ortópteros. Catálogo sistemático de toda la fauna de Filipinas, 2, 189–223.
Costa, A. (1864) Generi e specie d'insetti della fauna italiana. Annuario dell'Istituto e Museo di zoologia dell'Università di Napoli, 2, 128–138.
Deng, W.-A. (2016) Taxonomic study of Tetrigoidea from China. Huazhong Agricultural University, Wuhan, 341 pp.
Deng, W.-A., Zheng, Z. & Wei, S.-Z. (2007) Fauna of Tetrigoidea from Yunnan and Guangxi. Guangxi Science & Technology Press, Nanning, 458 pp.
Deng, W.-A., Zheng, Z. & Wei, S.-Z. (2009) A review of the genus Falconius Bolivar (Orthoptera: Tetrigoidea: Scelimeninae). Zootaxa, 1976, 63–68.
Deng, W.-A., Zheng, Z.-M., Wei, S.-Z. & Lin, M.-P. (2012) A systematic study of the genus Paragavialidium Zheng (Orthoptera: Tetrigoidea: Scelimeninae). Zootaxa, 3582, 48–56.
Ding, J.-H., Wen, T., Wu, X.-M., Boonmee, S., Eungwanichayapant, P.D. & Zha, L.S. (2017) Species diversity of Tetrigidae (Orthoptera) in Guizhou, China with description of two new species. Journal of Natural History, 51 (13–14), 741–760.
https://doi.org/10.1080/00222933.2017.1293749Fletcher, T.B. (1921) Catalogue of Indian Insects. Part I. Acrydidae (Tettigidae). Superintendent government printing, Calcutta, 40 pp.
Günther, K. (1935) Acrydiinen (Orth. Acrididae) aus dem mittleren Ostrorneo gesammelt von H. C. Siebers 1925. Arbeiten über Morphologische und Taxonomische Entomologie aus Berlin-Dahlem, 2 (4), 257–263, 7 figs.
Günther, K. (1937) Orthoptera celebica sarasiniana, Fam. Acrididae, Subfam. Acrydiinae. Treubia, 16, 165–195.
Günther, K. (1938a) Revision der Acrydiinae (Orth.) II. Scelimenae spuriae. Stettiner Entomologische Zeitung, 99, 117–148, 161–230.
Günther, K. (1938b) Revision der Acrydiinae, I. Sectiones Tripetalocerae, Discotettigiae, Lophotettigiae, Cleostrateae, Bufonidae, Cladonotae, Scelimenae verae. Mitteilungen aus dem Zoologischen Museum in Berlin, 23, 299–437.
Günther, K. (1938c) Acrydiinen (Orthoptera: Acrididae) von Neu Guinea hauptsächlich aus den Ausbeuten von Professor Dr. Bürgers (1912-13), Dr. E Mayr (1928), G. Stein (1931) and Miss L. E. Cheesman (1933-34). Nova Guinea, New Series, 2, 1–46.
Günther, K. (1939) Revision der Acrydiinae (Orthoptera), III. Sectio Amorphopi (Metrodorae Bol. 1887, aut.). Abhandlungen und Berichte aus den Staatlichen Museen für Tierkunde und Völkerkunde in Dresden, Series A (Zoology), Neue Folge, 20 (Bd. 1), 16–335.
Günther, K. (1941) Revision der Acrydiinenausbeute H. Sauters von Formosa (Orth.). Stettiner Entomologische Zeitung, 102, 145–165, 2 pls.
Günther, K. (1942) Die Acrydiinae der Oxforder Universitats-Expedition nach Sarawak vom Jahre 1932. Eos, Revista española de Entomología, 18, 339–346, 6 figs.
Günther, K. (1955) Über die Dornschrecken (Orth. Acrid. Tetrigidae) von Sumba und Flores mit faunenhistorischen Anmerkungen zur Verbreitung einiger Gattungsgruppen der Tetrigidae im südostasiatischen Inselbereich. Verhandlungen der Naturforschenden Gesellschaft in Basel, 66 (2), 147–175.
Gupta, S.K. & Chandra, K. (2017) A taxonomic study of the pygmy grasshopper (Orthoptera Tetrigidae) from India with description of a new species. Biodiversity Journal, 8 (2), 739–748
Gupta, S.K. & Chandra, K. (2018) A new species of genus Euscelimena Bolivar, 1887 (Orthoptera: Tetrigidae: Scelimeninae) from Central India. Munis Entomology & Zoology, 13 (1), 43–49.
Haan, W. (1843) Bijdragen tot de kennis der Orthoptera. Temminck [Ed.] Verhandelingen over de Natuurlijke Geschiedenis der Nederlansche Overzeesche Bezittingen, 19/20, 165–228.
Hancock, J.L. (1904) The Tettigidae of Ceylon. Spolia Zeylanica, 2, 97–157.
Hancock, J.L. (1906) Description of new genera and species of the orthopterous tribe Tettigidae. Entomological News, 17, 86–91.
Hancock, J.L. (1907a) Orthoptera Fam. Acridiidae. Subfam. Tetriginae. Genera Insectorum, 48, 1–79, 4 pls.
Hancock, J.L. (1907b) Studies of the Tetriginae (Orthoptera) in the Oxford University Museum. Transactions of the Entomological Society of London, 1907, 213–244, pl. 21.
Hancock, J.L. (1910) Notes on Ceylonese Tetriginae (Orthoptera), with description of some new species. Spolia Zeylanica, 6 (24), 140–149.
Hancock, J.L. (1913) Studies of Tetriginae (Acrydinae) from Sarawak Museum, Borneo. The Sarawak Museum Journal, 1 (3), 39–54.
Hancock, J.L. (1915) V. Indian Tetriginae (Acrydiinae). Records of the Indian Museum, 11, 55–137.
Hebard, M. (1930) Acrydiinae (Orthoptera, Acrididae) of Southern India. Revue Suisse de Zoologie, 36, 565–592.
https://doi.org/10.5962/bhl.part.117940Hollier, J. (2013) An annotated list of the Orthoptera (Insecta) species described by Henri de Saussure, with an account of the primary type material housed in the Muséum d’histoire naturelle de Genève. Part 4: The Acridomorpha excluding the superfamily Acridoidea. Revue Suisse de Zoologie, 120 (2), 203–219.
Hollier, J. (2016) The type specimens of Orthoptera (Insecta) species described by Ignacio Bolívar and deposited in the Muséum d'histoire naturelle de Genève. Revue Suisse de Zoologie, 123 (1), 21–33.
Hollier, J. & Heads, S. (2014) The type specimens of Orthoptera described by American entomologists in the collection of the Muséum d'histoire naturelle de Genève. Revue Suisse de Zoologie, 121 (1), 63–76.
Ichikawa, A. (1994) A revision of the family Tetrigidae (Orthoptera) of the Ryukyu Islands, southern Japan, with descriptions of new species and subspecies. (Part 1). Japanese Journal of Entomology, 62 (3), 457–474.
Jacobson, G.G. (1905 [1902–1905]) Orthoptera. In: Jacobson, G.G. & Bianchi, V.L. (Eds.), Orthopteroid and Pseudoneuropteroid Insects of Russian Empire and Adjacent Countries. A.F. Devrien Publication, St. Petersburg, pp. 6–466.
Jiang, G. & Zheng, Z. (1998) Grasshoppers and Locusts from Guangxi. Guangxi Normal University Press, Guilin, 390 pp.
Kevan, D.K.M. (1966) Some Orthoptera-Caelifera from the Philippine, Bismarck and Solomon Islands, with a few interesting records from New Guinea and the Moluccas. Entomologiske Meddelelser, 34, 375–420, 3 pls.
Kim, T.-W. & Pham, H.T. (2014) Checklist of Vietnamese Orthoptera (Saltatoria). Zootaxa, 3811 (1), 53–82.
https://doi.org/10.11646/zootaxa.3811.1.3Kirby, W.F. (1890) Incecta, excepting Coleoptera. In: Ridley, H.N., Notes on the zoology of Fernando Noronha. Zoological Journal of the Linnean Society, 20, pp. 530–548.
Kirby, W.F. (1910) A Synonymic Catalogue of Orthoptera (Orthoptera Saltatoria, Locustidae vel Acridiidae). The Trustees of the British Museum, London, 674 pp.
Kirby, W.F. (1914) Orthoptera (Acridiidae). In: Shipley, A.E. (Ed.), The Fauna of British India, Including Ceylon and Burma. Friedländer & Sohn, Berlin, pp. 1–176.
https://doi.org/10.5962/bhl.title.109305Liang, G.-Q. (2002) A new genus and a new species of Scelimenidae (Orthoptera: Tetrigoidea) [刺翼蚱科一新属新种(直翅目: 蚱总科)]. In: Zooscience. [动物科学]. Shaanxi Normal University Press, Xian, [陕西师范大学出版社, 西安], pp. 98–101.
Liang, G. & Zheng, Z. (1998) Orthoptera Tetrigoidea. Fauna Sinica, Insecta. Science Press Beijing, China. 278 pp.
Mahmood, K., Idris, A.B. & Salmah, Y. (2007) Tetrigidae (Orthoptera: Tetrigoidea) from Malaysia with the description of six new species. Acta Entomologica Sinica, 50 (12), 1272–1284.
Otte, D. (1979[1978]) The primary types of Orthoptera (Saltatoria, Blattodea, Phasmatodea, and Mantodea) at the Academy of Natural Sciences of Philadelphia. Proceedings of the Academy of Natural Sciences of Philadelphia, 130, 26–87.
Otte, D. (1997) Tetrigoidea and Tridactyloidea (Orthoptera: Caelifera) and Addenda to OSF Vols. 1-5. Orthoptera Species File, 6, 1–261.
París, M.G. (1994) Catálogo de tipos de ortopteroides (Insecta) de Ignacio Bolivar, I: Blattaria, Mantodea, Phasmoptera y Orthoptera (Stenopelmatoidea, Rhaphidophoroidea, Tettigonioidea, Grylloidea, Tetrigoidea). Eos, Revista española de Entomología, 69, 143–264.
Petersen, G. & Gaedike, H. (1970) Katalog der in den Sammlungen des Deutschen Entomologischen Institutes aufbewahrten Typen II. Beiträge zur Entomologie, 20 (1–2), 145–172.
Podgornaya, L.I. (1992) A new species of Tetrigidae (Orthoptera) from North Vietnam. Trudy Zoologicheskogo Instituta Akademii Nauk SSSR, 245, 46–49.
Rehn, J.A.G. (1904[1905]). Studies in the orthopterous subfamilies Acrydiinae (Tettiginae), Eumastacinae and Proscopinae. Proceedings of the Academy of Natural Sciences of Philadelphia, 56, 658–683.
Rehn, J. A. G. (1939) Studies in African Acrydiinae (Orthoptera, Acrididae). Part 2. New genera and species, and critical notes on previously-known forms. Proceedings of the Academy of Natural Sciences of Philadelphia, 90, 361–387.
Roonwal, M.L. (1979[1981]). Field bioecology and morphometry of some central Indian grasshoppers (Aridoidea), with notes on a swimming species (Tetrigoidea). Journal of the Zoological Society of India, 32 (1–2), 97–106.
Sandrasagara, T.R. (1949[1950]) A catalogue of the named Ceylonese Tetrigidae, Eumastacidae and Acrididae (Insecta: Orthoptera) in the collection of the Colombo Museum, with records of distribution. Records of the Indian Museum, 47, 135–157.
Saussure, H. (1862[1861]) Etudes sur quelques orthoptères du Musée de Genève. Annales de la Société Entomologique de France, 4 (1), 469–494.
Seeman, O.D., Lindquist, E.E. & Husband, R.W. (2018) A new tribe of tarsonemid mites (Trombidiformes: Heterostigmatina) parasitic on tetrigid grasshoppers (Orthoptera). Zootaxa, 4418 (1), 1–54.
https://doi.org/10.11646/zootaxa.4418.1.1Serville, G.A. (1838[1839]) Histoire naturelle des insectes. Orthoptères. Librairie Encyclopédique de Roret, Paris, xviii + 776 pp., 14 pls.
Shishodia, M.S. (1987) Tetrigidae: Orthoptera: Insecta (grouse locusts). Fauna of Orissa, 1, 91–102.
Shishodia, M.S. (1991) Taxonomy and zoogeography of the Tetrigidae (Orthoptera: Tetrigoidea) of north eastern India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 140, 1–204.
Shishodia, M.S. (1995) Insecta: Orthoptera: Tetrigoidea. State Fauna Series 4 Fauna of Meghalaya, 3, 165–208.
Shishodia, M.S. (2007) Insecta: Orthoptera: Tetrigidae. Fauna of Mizoram. State (Mizoram) Fauna Series, Zoological Survey of India, 14, 217–229.
Shishodia, M.S. & Hazra, A.K. (1985) Insecta: Orthoptera. Records of the Zoological Survey of India, Calcutta, 82 (1–4), 15–32.
Shishodia, M.S., Chandra, K. & Gupta, S.K. (2010) An annotated checklist of Orthoptera (Insecta) from India. Records of the Zoological Survey of India, Miscellaneous Publication, Occasional Paper, 314, 1–366.
Skejo, J. (2017) Taxonomic Revision of the Pygmy Devils (Tetrigidae: Discotettiginae) with Online Social Media as a New Tool for Discovering Hidden Diversity. Diploma thesis, Faculty of Science, University of Zagreb, Zagreb, 246 pp.
Skejo, J. & Bertner, P. (2017) No more dust and exoskeletons—in vivo photographic records provide new data on Eufalconius pendleburyi Günther, 1938 (Orthoptera: Tetrigidae) from the Titiwangsa Mts. Annales Zoologici, 67 (4), 665–672.
https://doi.org/10.3161/00034541ANZ2017.67.4.003Skejo, J. & Caballero, J.H.S. (2016) A hidden pygmy devil from the Philippines: Arulenus miae sp. nov.—a new species serendipitously discovered in an amateur Facebook post (Tetrigidae: Discotettiginae). Zootaxa, 4067 (3), 393–393.
http://doi.org/10.11646/zootaxa.4067.3.7Skejo, J., Gupta, S.K. & Tumbrinck, J. (2018) Nymph inadvertently described as new species for a fourth time? On the identity of Euscelimena hardi (Tetrigidae: Scelimeninae) with general remarks on the identification of pygmy grasshopper nymphs. Zootaxa, 4418 (1), 93–97.
https://doi.org/10.11646/zootaxa.4418.1.6Stål, C. (1877) Orthoptera nova ex Insulis Philippinis descripsit. Öfversigt af Kongliga Vetenskaps-Akademiens Förhandlinger, 34 (10), 33–58.
Steinmann, H. (1970) Check-list of the Tetricidae (Orthoptera) of the Oriental faunal region. Acta Zoologica Academiae Scientiarum Hungaricae, 16, 215–240
Stoll, C. (1813) Représentation exactement colorée d'après nature des spectres ou phasmes, des mantes, des sauterelles, des grillons, des criquets et des blattes. Jan Christiaan Sepp, Amsterdam, 79 pp.
Storozhenko, S.Y. (2014) A new species of the genus Falconius Bolívar, 1898 (Orthoptera: Tetrigidae, Scelimeninae) from Vietnam. Entomological Review, 94 (4), 602–604.
https://doi.org/10.1134/S0013873814040125Storozhenko, S.Y. (2017) A new species of the genus Falconius Bolívar, 1898 (Orthoptera: Tetrigidae, Scelimeninae) from Cambodia. Far Eastern Entomologist, 332, 1–6.
https://doi.org/10.25221/fee.344.1Storozhenko, S.Y. & Dawwrueng, P. (2014) Three new species of the subfamily Cladonotinae (Orthoptera: Tetrigidae) from Thailand. Zootaxa, 3811 (3), 325–337.
https://doi.org/10.11646/zootaxa.3811.3.3Storozhenko, S.Y. & Dawwrueng, P. (2015) New and little-known pygmy grasshoppers (Orthoptera: Tetrigidae) from Thailand. Zootaxa, 4052 (5), 527–554.
https://doi.org/10.11646/zootaxa.4052.5.2Storozhenko, S.Y. & Paik, J.C. (2011) Review of the genus Bidentatettix (Zheng, 1992) (Orthoptera: Tetrigidae, Cladonotinae). Korean Journal of Soil Zoology, 15 (1–2), 48–52.
Storozhenko, S.Y. & Pushkar, T.I. (2017) A new genus of pygmy locusts (Orthoptera: Tetrigidae: Cladonotinae) from the Malay peninsula. Annales Zoologici, 67 (1), 47–53.
https://doi.org/10.3161/00034541ANZ2017.67.1.006Tan, M.K. & Artchawakom, T. (2015) A new species from the genus Gorochovitettix (Tetrigidae: Metrodorinae) from Thailand. Zootaxa, 3990 (3), 444–450.
https://doi.org/10.11646/zootaxa.3990.3.9Tan, M.K., Choi, J. & Shankar, N. (2017) Trends in new species discovery of Orthoptera (Insecta) from Southeast Asia. Zootaxa, 4238 (1), 127–134.
https://doi.org/10.11646/zootaxa.4238.1.10Tan, M.K., Storozhenko, S.Y., Hwang, W.S. & Meier, R. (2017) Integrative taxonomy reveals two sympatric species of the genus Eucriotettix Hebard, 1930 (Orthoptera: Tetrigidae). Zootaxa, 4268 (3), 377–394.
https://doi.org/10.11646/zootaxa.4268.3.4Tan, M.K. & Wahab, R.A. (2018) Preliminary study on the diversity of Orthoptera from Kuala Belalong Field Studies Centre, Brunei Darussalam, Borneo. Journal of Orthoptera Research, 27 (2), 119–142.
https://doi.org/10.3897/jor.27.24152Tinkham, E.R. (1936) A new species of Eugavialidium from Formosa (Orth. Acrid. Acridinae). Transactions of the Natural History Society of Formosa, 26, 207–210.
Tinkham, E.R. (1937) Notes on the identity of Formosan Acrydiinae with descriptions of a new genus and two new species (Orth.: Acrid.). Transactions of the Natural History Society of Formosa, 27, 229–243, 1 pl.
Tumbrinck, J. (2014) Taxonomic revision of the Cladonotinae (Orthoptera: Tetrigidae) from the islands of South-East Asia and from Australia, with general remarks to the classification and morphology of the Tetrigidae and descriptions of new genera and species from New Guinea and New Caledonia. In: Telnov, D. (Ed.), Biodiversity, Biogeography and Nature Conservation in Wallacea and New Guinea, II, pp. 350–396.
Tumbrinck, J. (2018) A new species of Thoradonta from New Guinea with some remarks on other Tetrigidae (Orthoptera) taxa from Indo-Australia. Suara Serangga Papua (SUGAPA digital), 11 (1), 27–46.
https://doi.org/10.19269/sugapa2018.11(1).05Uvarov, B.P. (1927) Distributional records of Indian Acrididae. Records of the Indian Museum, 29, 233–239.
Vanschuytbroeck, P. (1980) Catalogue et liste du matériel typique des Tetrigidae conservés dans les collections entomologiques de l'Institut royal des Sciences naturelles de Belgique. Bulletin de l'Institut Royal des Sciences Naturelles de Belgique, 52 (22), 1–25.
Walker, F. (1871) Catalogue of the Specimens of Dermaptera Saltatoria in the Collection of the British Museum. Trustees of the British Museum, 5, 811–850.
Willemse, C. (1928) Revision des Acridiodea, décrites par De Haan, avec descriptions de nouvelles espèces. Zoologische Mededelingen, Leiden, 11 (1), 1–27.
Willemse, C. (1930) Fauna Sumatrensis (Bijdrage Nr. 62): Preliminary revision of the Acrididae (Orthoptera). Overgedrukt uit het Tijdschrift voor Entomologie, 73, 1–210.
Yin, X.-C., Shi, J. & Yin, Z. (1996) Synonymic Catalogue of Grasshoppers and their Allies of the World (Orthoptera: Caelifera). China Forestry Publishing Hous, Beijing, 1266 pp.
Zha, L., Yu, F., Boonmee, S., Eungwanichayapant, P.D. & Wen, T. (2017) A new species of the genus Scelimena (Orthoptera: Tetrigidae) in Thailand and notes on its biology and ecology. Entomological News, 126 (5), 372–382.
https://doi.org/10.3157/021.126.0505Zha, L.-S., Wen, T.-C., Pan, Z.-X. & Hyde, K.D. (2016) Taxonomy of Paragavialidium (Orthoptera: Tetrigidae: Scelimeninae) with description of one new species and notes on ecology and habits. Entomological News, 126 (1), 43–51.
https://doi.org/10.3157/021.126.0106Zheng, Z.M. (1992) Orthoptera: Tetrigidae. In: Huang, F. (Ed.), Insects of Wuling Mountains Area, Southwestern China. Science Press, Beijing, pp. 81–86.
Zheng, Z.M. (1994) A new genus and three new species of Scelimenidae from China (Orthoptera: Terigoidea [Tetrigoidea]). Journal of Hubei University (Natural Science), 16 (1), 1–5.
Zheng, Z.M. (2004) Orthoptera: Tetrigoidea. In: Insects from Mt. Shiwandashan Area of Guangxi. Chinese Forestry, Beijing, pp. 69–89.
Zheng, Z.M. (2005) Fauna of Tetrigoidea from Western China. Science Press, Beijing, 501 pp. [in Chinese with English summary]
Zheng, Z.M., Wang B., Li, P. & Niu, N. (2006) A taxonomic study on the genus Falconius Bolivar (Orthoptera, Scelimenidae) from China. Acta Zootaxonomica Sinica, 31, 821–823.
Zheng, Z.M. (2014) Key to the species of Falconius, Hedotettix and Euparatettix (Orthoptera: Tetrigoidea) from China with description of two new species. Journal of Shangqiu Normal University, 30 (3), 1–14.
Zheng, Z.M. & Cao, C. (2011) A review of the genus Paragavialidium Zheng (Orthoptera, Scelimenidae) with descriptions of two new species. Acta Zootaxonomica Sinica, 36 (3), 742–745.
Zheng, Z.M. & Jiang, G. (1997) Two new species of Scelimenidae and first description of female of Bolivaritettix longzhouensis (Orthoptera: Tetrigoidea) from China. Entomotaxonomia, 19 (3), 164–168.
Zheng, Z.M. & Liang, G. (1985) Studies on the Tetrigidae from China. Entomotaxonomia, 7 (1), 51–56.
Zheng, Z.M. & Ou, X. (2003) Four new species of Scelimenidae from Yunnan Province (Orthoptera, Tetrigoidea). Acta Zootaxonomica Sinica, 28 (4), 673–678.
Zheng, Z.M. & Yang, R. (2015) A revision of the genus Bolivaritettix Gunther (Tetrigoidea: Metrodoridae) with descriptions of three new species in China. Journal of Shangqiu Normal University, 31 (3), 1–18.